Photoassimilates are loaded along the entire phloem transport pathway, from photosynthetic leaves to importing sinks. While most photoassimilate loading occurs in photosynthetically active leaves, root-produced metabolites, such as amino acids, move readily from xylem to phloem particularly at the stem nodes. Phloem loading also occurs in storage organs during periods when reserves are remobilised and exported. Indeed, the membrane transport events contributing to phloem loading were first examined using export of sucrose remobilised from the endosperm of germinating castor bean seed as an experimental model (Kriedemann and Beevers 1967).
This section focuses on phloem loading in the leaves. It analyses the cellular pathways for assimilate loading, and the regulatory controls.